Mutual Aid; a factor of evolution

ns, etc. Societies of rodents; of ruminants; of monkeys. Mutual Aid in the struggle for life. Darwin's arguments to prove the struggle for lif

an cliff, and each of the lakes and ponds with which Northern America, Northern Europe, and Northern Asia are dotted tell us at that time of the year the tale of what mutual aid means for the birds; what force, energy, and protection it confers to every living being, however feeble an

of plovers and sand-coursers run over the beach, searching their. food, whistling, and simply enjoying life. Further on, on alm

d terns set to chase the robber. Maddened by hunger, the robber soon abandons his usual precautions: he suddenly dashes into the living mass; but, attacked from all sides, he again is compelled to retreat. From sheer despair he falls upon the wild ducks; but the intelligent, social birds rapidly gather in a flock and fly away if the robber is an erne; they plunge into the lake if it is a falcon; o

in the Arctic

to a height of from two to five hundred feet, literally covered with sea-birds, whose white breasts show against the dar

there, the extremely sociable kittiwake-gulls, among whom quarrels are rare and short; the prepossessing polar guillemots, which continually caress each other; the egoist she-goose, who has repudiated the orphans of a killed comrade; and, by her side, another female who adopts any one's orphans, and now paddles surrounded by fifty or sixty youngsters, whom she conducts and cares for as if they all were her own breed. Side by side with the penguins, which steal one another's eggs, you have the dotterels, whose family relations are so "charming and touching" that even passionate hunters r

t charming descriptions of the peace and harmony which prevail in almost all these nesting associations. As to the protection derived by the weakest birds from their unions, it is evident. That excellent observer, Dr. Coues, saw, for instance, the little cliff-swallows nesting in the immediate neighbourhood of the prairie falcon (Falco polyargus). Th

ng nuthatchers (Sitta caesia), together with tit-mouses, chaffinches, wrens, tree-creepers, or some wood-peckers.5 In Spain the swallow is met with in company with kestrels, fly-catchers, and even pigeons. In the Far West of America the young horned larks live in large societies, together with another lark (Sprague's), the skylark, the Savannah sparrow, and several specie

efore they start, and they evidently discuss the particulars of the journey. Some species will indulge every afternoon in flights preparatory to the long passage. All wait for their tardy congeners, and finally they start in a certain well chosen direction - a fruit of accumulated collective experience - the strongest flying at t

ich one sees in the north, after the birds have arrived at their breeding-places on the Yenisei or in the northern counties of England. For many days in succession - sometimes one month - they will come together every morning for one hour, before flying in search of food - perhaps discussing the spot where they are going to build their nests.8 And if, during the migration, their columns are overtaken by a storm, birds of the most different species will be brough

of which are sociable animals. When the Europeans came to settle in America, they found it so densely peopled with buffaloes, that pioneers had to stop their advance when a column of migrating buffaloes came to cross the route they followed; the march past of the dense column lasting sometimes for two and three days. And when the Russians took

r north the reindeer aggregate in numberless herds; while still further north we find the herds of the musk-oxen and numberless bands of polar foxes. The coasts of the ocean are enlivened by flocks of seals and morses; its waters, by shoals of sociable cetaceans; and even in the depths of the great plateau of Central Asia we find herds of wild horses, wild donkeys, wild camels, and wild sheep. All these mammals live in societies and nations sometimes numbering hundreds of thousands of individ

the thirties, also saw the Labrador wolves hunting in packs, and one pack following a man to his cabin, and killing the dogs. During severe winters the packs of wolves grow so numerous as to become a danger for human settlements, as was the case in France some five-and-forty years ago. In the Russian Steppes they never attack the horses otherwise than in packs; and yet they have to sustain bitter fights, during which the horses (according to Kohl's testimony) sometimes assume offensive warfare, and in such cases, if the wolves do not retreat promptly, they run the risk of being surrounded by the horses and killed by their hoofs. The prairie-wolves (Canis latrans) are known to associate in bands of from twenty to thirty individuals when they chase a buffalo occasionally separated from its herd.12 Jackals, which are most courageous and may be considered as one of the most intelligent representatives of the dog tribe, always hunt in packs; thus united, they have no fear of the bigger carnivores.13 As to the wild dogs of Asia (the Kholzuns, or Dholes), Williamson saw their large packs attacking all larger animals save elephants and r

n a close intercourse, and when the pine-cones become rare in the forest they inhabit, they emigrate in bands. As to the black squirrels of the Far West, they are eminently sociable. Apart from the few hours given every day to foraging, they spend their lives in playing in numerous parties. And when they multiply too rapidly in a region, they assemble in bands, almost as numerous as those of locusts, and move southwards, devastating the forests, the fields, and the gardens; while foxes, polecats, falcons, and nocturnal birds of prey follow their thick column

ly returning to his citizen's duties - he begins inventing the most diabolic means for the extermination of the little robbers. All kinds of rapacious birds and beasts of prey having proved powerless, the last word of science in this warfare is the inoculation of cholera! The villages of the prairie-dogs in America are one of the loveliest sights. As far as the eye can embrace the prairie, it sees heaps of earth, and on each of them a prairie-dog stands, engaged in a lively conversation with its neighbours by means of short barkings. As soon as the approach of man is signalled, all plunge in a moment into their dwellings; all have disappeared as by enchantment. But if the danger is over, the little creatures soon reappear. Whole families come out of their galleries and indulge in play. The young ones scratch one

ne with other species, and they have attained a very high degree of intellectual development. In their villages, always disposed on the shores of lakes and rivers, they take into account the changing level of water; their domeshaped houses, which are built of beaten clay interwoven with reeds, have separate corners for organic refuse, and their halls are well carpeted at winter time; they are warm, and, nevertheless, well ventilated. As to the beavers, which are endowed, as known, with a most sympathetic character, t

o not gather in societies for life in common, and which are not even endowed with intense parental feelings, cannot live without coming together for play. Dietrich de Winckell, who is considered to be among the best acquainted with the habits of hares, describes them as passionate players, becoming so intoxicated by their play that a hare has been known to take an approaching fox for a playmate.16 As to the rabbit, it lives in societies, and its family life is entirely built upon the image of the old patriar

e of the earth were it not for their sociable spirit. When a beast of prey approaches them, several studs unite at once; they repulse the beast and sometimes chase it: and neither the wolf nor the bear, not even the lion, can capture a horse or even a zebra as long as they are not detached from the herd. When a drought is burning the grass in the prairies, they gather in herds of sometimes 10,000 individuals strong, and migrate. And when a snow-storm rages in the Steppes, each stud keeps close together, and repairs to a protected ravine. But if confidence disappears, or

he Amur, I could ascertain how thinly-peopled with fallow deer these mostly uninhabited regions are.19 Two years later I was travelling up the Amur, and by the end of October reached the lower end of that picturesque gorge which the Amur pierces in the Dousse-alin (Little Khingan) before it enters the lowlands where it joins the Sungari. I found the Cossacks in the villages of that gorge in the greatest excitement, because thousands and thousands of fallow deer were crossing the Amur where it is narrowest, in order to reach the lowlands. For several days in succession, upon a length of some forty miles up the river, the Cossacks were butchering the deer as they crossed the Amur, in which already floated a good deal of ice. Thousands were killed every day, and the exodus nevertheless continued. Like migrations were never seen either before or since, and this one must have been called for by an early and heavy snow-fall in the Great Khingan, which compelled the deer to make a desperate attempt at reaching the lowlands in the east of the Dousse mountai

the wild boars, and find a word of praise for their powers of association in the case of an attack by a beast of prey.21 The hippopotamus and the rhinoceros, too, would occupy a place in a work devoted to animal sociability. Several striking pages might be given to the sociability and mutual attachment of t

nd so on - are sociable in the highest degree. They live in great bands, and even join with other species than their own. Most of them become quite unhappy when solitary. The cries of distress of each one of the band immediately bring together the whole of the band, and they boldly repulse the attacks of most carnivores and birds of prey. Even eagles do not dare attack them. They plunder our fields always in bands - the old ones taking care for the safety of the commonwealth. The little tee-tees, whose childish sweet faces so much struck Humboldt, embrace and protect one another when it rains, rolling their tails over the necks of their shivering comrades. Several species display the greatest solicitude for their wounded, and do not abandon a wounded comrade during a retreat till they have ascertained that it is dead and that they are helpless to restore it to life. Thus James Forbes narrated in his Oriental Memoirs a fact of such resistance in reclaiming from his hunting party the dead body of a female monkey that one fully understands why "the witnesses of this extraordinary scene resolved never again to

only, are relatively few, and their numbers are limited. Nay, it appears very probable that, apart from a few exceptions, those birds and mammals which are not gregarious now, were living in societies before man multiplied on the earth and waged a permanent war agai

ls combine, under the pressure of exceptional circumstances, to emigrate. In this last case, it becomes a voluntary deviation from habitual moods of life. The combination sometimes appears in two or more degrees - the family first, then the group, and finally the association of groups, habitually scattered, but uniting in case of need, as we saw it with the bisons and other ruminants. It also takes higher forms, guaranteeing more independence to the individual without depriving it of the benefits of social life. With most rodents the individual has its own dwelling, which it can retire to when it prefers being left alone; but the dwellings are laid out in villages and cities, so as to guar

, are, together with dancing and singing, mere manifestations of an excess of forces -"the joy of life," and a desire to communicate in some way or another with other individuals of the same or of other species - in short, a manifestation of sociability proper, which is a distinctive feature of all the animal world.26 Whether the feeling be fear, experienced at the appearance of a bird of prey, or "a fit of gladness" which bursts out when the animals are in good health and especially when young, or merely the desire of giving play to an excess of impressions and of vital power - the necessity of communicating impressio

f Man (ch. xiii). Visitors of the London Zoological Gardens also know the bower of the satin bower-bird. But this habit of dancing seems to be much more widely spread than was formerly believed, and Mr. W. Hu

Chauna chavarris), to which the English have given the most unimaginative misnomer of "crested screamer." These birds sometimes assemble in immense flocks, and in such cases

lock took up the strains, and after it the next, and so on, until once more the notes of the flocks on the opposite shore came floating strong and

of birds covering the marsh for miles around burst forth in a tremendous evening song. . . . It was a concert well worth riding a hundred miles to hear."27 It may be added that like all sociable animals, the chakar easily

ance to hunger and cold, which are mentioned by Darwin and Wallace, are so many qualities making the individual, or the species, the fittest under certain circumstances, we maintain that under any circumstances sociability is the greatest advantage in the struggle for life. Those species which willingly or unwillingly abandon it are doomed to decay; while those animals which know best how to combine, have the greatest chances of survival and of further evolution, although they may be inferior to others in each of the faculties enumerated by Darwin and Wallace, save the intellectual faculty. The highest vertebrates, and especially mankind, are the best proof of this assertion. As to the intellectual faculty, while every Darwinist will agree with Dar

irs to rest, and from which it never deviates; and so on.28 We have any numbers of direct observations of the peace that prevails in the nesting associations of birds, the villages of the rodents, and the herds of grass-eaters; while, on the other side, we know of few sociable animals which so continually quarrel as the rats in our cellars do, or as the morses, which fight for the possession of a sunny place on the shore. Sociability thus puts a limit to physical struggle, and leaves room for the development of better moral feelings. The high development of parental love in all classes of animals, even with lions and tigers, is generally known. As to the young birds and mammals whom we continually see associating, sympathy - not love - attains a further development in their associations. Leaving aside the really touching facts of mutual attachment and compassion which have been recorded as regards domesticated animals and with animals kept in captivity, we have a number of well certified facts of compassion between wild animals at liberty. Max

of our century. Life is struggle; and in that struggle the fittest survive. But the answers to the questions, "By which arms is this struggle chiefly carried on?" and "Who are the fittest in the struggle?" will widely differ according to the importance given to the two different aspects of the struggle: the direct one, for food and safety among separate individuals, and the struggle which Darwin described as "metapho

o not find them sufficiently convincing. If we refer to the paragraph entitled "Struggle for Life most severe between Individuals and Varieties of the same Species," we find in it none of that wealth of proofs and illustrations which we are accustomed to find in whatever Darwin wrote. The struggle between individuals of the same species is not illustrated under that heading by even one single instance: it

ich fill nearly the same place in nature; but probably in no case could we precisel

fe between closely-allied Animals and Plants often most severe"), he makes the following rem

ecessary, and there may be cases in which the weaker species, physically, may prevail by its power of more rapid multipli

the assumption based? The same remark must be made concerning the indirect argument in favour of a severe competition and struggle for life within each species, which may be derived from the "extermination of transitional varieties," so often mentioned by Darwin. It is known that for a long time Darwin was worried by the difficulty which he saw in the absence of a long chain of intermediate forms between closely-allied species, and that he found the solution of this difficulty in the supposed extermination of the intermediate forms.33 However, an attentive reading of the different chapters in which Darwin and Wallace speak of this subject soon brings one to the conclusion that the word "extermination" does not mean real extermination; the same remark which Darwin made concerning his expression: "struggle for existence," evidently applies to the word "extermination" as well. It can by no means be understood in its direct sense, but must be taken "in its metaphoric sense." If we start from the supposi

e with the slow snail, as with the swift bird; physical changes are continually going on in every given area; and new varieties among animals consist in an immense number of cases-perhaps in the majority - not in the growth of new weapons for snatching the food from the mouth of its congeners - food is only one out of a hundred of various conditions of existence - but, as Wallace himself shows in a charming paragraph on the "divergence of characters" (Darwinism, p. 107), in forming new habits, moving to new abodes, and taking to new sorts of food. In all such cases there will be

action of these agencies. It is known that portions of a given species will often take to a new sort of food. The squirrels, for instance, when there is a scarcity of cones in the larch forests, remove to the fir-tree forests, and this change of food has certain well-known physiological effects on the squirrels. If this change of habits does not last - if next year the cones are again plentiful in the dark larch woods - no new variety of squirrels will evidently arise from this cause. But if part of the wide area occupied by the squirrels begins to have its physical characters altered - in consequence of, let us say, a milder climate or desiccation, which both bring about an increase of the pine forests in proportion

d and New World, returning, in all probability, after a time to the pastures which they had, in the course of their migrations, formerly left.34 Consequently, if we do not find now, in Asia, all the intermediate links between the present wild horse and its Asiatic Post–Tertiary ancestors, this does not mean at all that the intermediate l

it must be used in its metaphoric sense. As to "competition," this expression, too, is continually used by Darwin (see, for instance, the paragraph "On Extinction") as an image, or as a way-of-speaking, rather th

of existence continually going on within every animal species is - to use Pro

m year to year the population remained stationary, for the simple reason that one-third of the new-born died before reaching their sixth month of life; one-half died within the next four years, and out of each hundred born, only seventeen or so reached the age of twenty. The new-comers went away before having grown to be competitors. It is evident that if such is the case with men, it is still more the case with animals. In the feathered world the destr

y of the region, but by what it is every year under the most unfavourable conditions. So that, for that reason alone, competition hardly can be a normal condition. but other causes intervene as well to cut, down the animal population below even that low standard. If we take the horses and cattle which are grazing all the winter through in the Steppes of Transbaikalia, we find them very lean and exhausted at the end of the winter. But they grow exhausted not because there is not enough food for all of them - the grass buried under a thin sheet of snow is everywhere in abundance - but because of the difficulty of getting it from beneath the snow, and this difficulty is the same for all horses alike. Besides, days of glazed frost are common in early spring, and if several such days come in succession the horses grow still more exhausted. But then comes a snow-storm, which compels the already weakened animals to remain without any food for several days, and very great numbers of them die. The los

amidst a nature which might support a hundred times as many ants as are actually living. Dr. Altum, a German forester, who wrote a very interesting book about animals injurious to our forests, also gives many facts showing the immense importance of natural checks. He says, that a succession of gales or cold and damp weather during the exodus of the pine-moth (Bombyx pini) destroy it to incredible amounts, and during the spring of 1871 all these moths disappeared at once, probably killed by a succession of cold nights.36 Many like examples relative to various insects could be quoted from various parts of Europe. Dr. Altum also mentions the bird-enemies of the pine-moth, and the immense amount of its eggs destroyed by foxes; but he adds that the parasitic fungi which periodically infest it are a far more terrible enemy than any bird, because

ors, as they are in Siberia; and Ch. Dixon saw the red grouse so pressed during some exceptionally severe winters, that they quitted the moors i

rs, even with the most rapidly-multiply ing animals. Thus, some sixty years ago, the sousliks suddenly disappeared in the neighbourhood of Sarepta, in South–Eastern Russ

he rule in the animal world. Those who survive a famine, or a severe epidemic of cholera, or small-pox, or diphtheria, such as we see them in uncivilized countries, are neither the strongest, nor the healthiest, nor the most intelligent. No progress could be based on those survivals - the less so as all survivors usually come out of the ordeal with an impaired health, like the Transbaikalian horses just mentioned, or the Arctic crews, or the garrison of a fortress which has been compelled to live for a few months on half rations, and comes out of its experience with a broken health, and subsequently shows a quite abnormal mortality. All that natural selection can do in times of calamities is to spare the individuals e

s, they rear their cattle - and thus avoid competition; and natural selection picks out of the ants' family the species which know best how to avoid competition, with its unavoidably deleterious consequences. Most of our birds slowly move southwards as the winter comes, or gather in numberless societies and undertake long journeys - and thus avoid competition. Many rodents fall asleep when the time comes that competition should set in; while other rodents store food for the winter, and gather in large villages for obtaining the necessary protection when at work. The reindeer, when the liche

"Therefore combine - practise mutual aid! That is the surest means for giving to each and to all the greatest safety, the best guarantee of existence and progress, bodily, intellectual, and moral." That is what Nature teaches us; and that is what all those animals which have attai