Darwiniana

ecurring tendency to vary--to vary to a greater or to a less extent. Such a variety, I pointed out to you, might arise from causes which we do not understand; we the

thout any exercise of that process which we called selection. And then I stated further, that by such selection, when exercised artificially--if you took care to breed only f

an be produced by this process of selective breeding? In considering this question, it will be useful to class the characteristics, in respec

t there might be an absolute difference in the number of the vertebrae of the back, as in the sacral vertebras of the Pouter; and so great is the extent of the variation in these and similar characters that I pointed out to you, by reference to the skeletons and the diagrams, that these extreme varieties may absolutely differ more from one another in their structural characters than do what naturalists call distinct SPECIES of pigeons; that is to say, that they differ so much in

bly use it a good deal, I had better perhaps devo

ROVINCES; and so on from a PROVINCE to a CLASS, from a CLASS to an ORDER, from ORDERS to FAMILIES, and from these to GENERA, until we come at length to the smallest groups of ani

iarities, then all naturalists agree in calling them two species; that is what is meant by the use of the word species--that is to say, it is, for the practical naturalist, a mere question of structural differences. [*] We have seen now--to repeat this point once

s." Whether a physiological test between species exist or n

gical differences of varieties amount in degree to those observed between forms which

teristics, there is no doubt that they are capable of

siological peculiarities--as the peculiar habit of tumbling, in the Tumbler--the peculiarities of flight, in the "homing" birds--the strange habit of spreading out the tail, and walking in a peculiar fashion, in the Fantail--and

you do find a check, and that is in the reproductive processes. For there is a most singular circumstance in respect to natural species--at least about some of them--and it would be sufficient for the purposes of this argument if it were true of only one of them, but there is, in fact, a great number of such cases--and that is, that, similar as they may appear to be to mere races or breeds, they present a marked peculiarity in the reproductive process. If you breed from the male and female of the same race, you of course have offspring of the like kind, and if you make the offspring breed together, you obtain the same result, and if you breed from these again, you will sti

tive elements necessary to generation. It is said to be invariably the case with the male mule, the cross between the Ass and the Mare; and hence it is, that, although crossing the Horse with the Ass is easy enough, and is constantly done, a

ing of one species with another, but you see it does not in reality do anything of the kind. There is nothing in this fact that hybrids cannot breed with each other, to establish such a theory; there is

s between distinct species. The mongrels are, so far as we know, fertile with one another. But between species, in many cases, you cannot s

en the Fantail and the Pouter, the Carrier and the Tumbler, or any other variety or race you may name--so far as we know at present--there is no difficulty in breeding together the mongrels. Take the Carrier and the Fantail, for instance, and let them represent the Horse and the Ass in the case of distinct species; then you have, as the result of their breeding, the Carrier-Fantail mongrel,--we will say the m

rather closely, I think you will find that it is not worth very much. The first question of course is, Do they thus return to the primitive stock? And commonly as the thing is assumed and accepted, it is extremely difficult to get anything like good evidence of it. It is constantly said, for example, that if domesticated Horses are turned wild, as they have been in some parts of Asia Minor and South America, that they return at once to the primitive stock from which they were bred. But the first answer that you make to this assumption is, to ask who knows what the primitive stock was; and the second answer is, that in that case the wild Horses of Asia Minor ought to be exactly like the wild Horses of South America. If they are both

he purpose of bringing the animal from its wild to its domesticated state. I do not see anything very wonderful in the fact, if it took all that trouble to get it from a wild state, that it should go back into its original state as soon as you removed the conditions which produced the variation to the domesticated form. There is an important fact, however, forcibly brought forward by Mr. Darwin, which has been noti

me ago, in a baker's cart, near Rothesay, in Bute: it had the long stripe down the back, and stripes on the shoulders and legs, just like those of the Ass, the Quagga, and the Zebra. Now, if we interpret the theory of recurrence as applied to this case, might it not be said that here was a case of a variation exhibiting the characters and conditions of an animal occupying something like an intermediate position between the Horse, the Ass, the Quagga, and the Zebra, and from which these had been developed? In the same way with regard even to Man. Every anatomist will tell you that there is nothing commoner, in dissecting the human body, than to meet with what are called muscular variations--that is, if you dissect two bodies very

is concerned; and in favour of a physiological limitation. By selective breeding we can produce structural divergences as

occurred in domesticated animals. I told you, that almost always the origin of these varieties is overlooked, so that I could only produce two or three cases, as that of Gratio Kelleia and of the Anconsheep. People forget, or do not take notice of them until they come to have a prominence; and if that is true of artificial cases, under our own eyes, and in animals in our own care, how much more difficult it must be to have at first hand good evidence of the origin of varieties in nature! Indeed, I do not know that it is possible by direct evidence to prove

f every naturalist, and of any person who has ever turned any attention at all to the characteristics of plants

ach have peculiar characteristics of colour and physiognomy; but you must recollect that the characters of these races go very far deeper--they extend to the bony structure, and to the characters of that most important of all organs to us--the brain; so that, among men belonging to different races, or even within the same race, one man shall have a brain a third, or half, or even seventy per cent, bigger than another; and if you take the whole range of human brains, you will find a variation in some cases of a hundred per cent. Apart from these variations in the size of the brain, the characters of the skull vary. Thus if I draw t

being tigers. The Siberian tiger has a thick fur, a small mane, and a longitudinal stripe down the back, while the tigers of Java and Sumatra differ in many important respects from the tigers of Northern Asia. So lions vary; so birds vary; and so, if you go further back and lower down in creation, you find that fishes vary. In different streams, in the same country even

war about it; some of them wanting to make out that there are many species of it, and others maintaining that they

ve described--as spontaneous varieties--and that those varieties may be perpetuated in the same way that I have shown you spontaneo

ve that, at present, I say nothing about species; I wish to confine myself to the consideration of the production of those natural races which everybody admits to exist. The quest

is FOOD, by which I mean food in the broadest sense, the supply of the materials necessary to the existence of an organic being; in the case of a plant the inorganic matters, such as carbonic acid, water, ammonia, and the earthy salts or salines; in the case of the animal the inorganic and organic matters, which we have seen they require; then these are all, at least the first two, what we may call the inorganic or physical conditions of existence. Food takes a mid-place, and then come the organic conditions; by which I mean the conditions which depend upon the state of the rest of the organic creation, upon the number and kind of living beings, with which an animal is surrounded. You may class these under two heads: there are organic beings, which operate as opponents, and there are organic beings which operate as helpers to any given organic creature. The opponents may be of two kinds: there are the indirect opponents, which are what we may call rivals; and there are the direct opponents, those which strive to destroy the creature; and these we call enemies.

st notion of properly estimating them until the publication of Mr. Darwin's work, which has placed them before us with remarkable clearness; and I must endeavour, as far as

ant, with no opponents, no helpers, and no rivals; it is to be a "fair field, and no favour." Now, I will ask you to imagine further that it shall be a plant which shall produce every year fifty seeds, which is a very moderate number for a plant to produce; and that, by the action of the winds and currents, these seeds shall be equally and gradually distributed over the whole surface of the land. I want you now to trace out what will occur, and you will observe that I am not talking fallaciously any more than a mathematician does when he expounds his problem. If you show that the conditions of your problem are such as may actually occur in Nature and do not tr

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the total number really is; but, at any rate, you will understand the meaning of all those noughts. Then you see that, at the bottom, I have taken the 51,000,000 of square miles, constituting the surface of the dry land; and as the number of square feet are placed under and subtracted from the number of seeds that would be produced in th

ve never yet been disproved and never will be--he showed that in consequence of the increase in the number of organic beings in a geometrical ratio, while the means of existence cannot be made to increase in the same ratio, that there must come a time when the number of organic beings will be in excess of the power of production of nutr

, as it were, forty-nine chances to one against its growing up; it depends upon the most fortuitous circumstances whether any one of these fifty seeds shall grow up and flourish, or whether it shall die and peri

, that the smallest possible start given to any one seed may give it an advantage which will enable it to get ahead of all the others; anything that will enable any one of these seeds to germinate six hours before any of the others will, other things being alike, enable it to choke them out altogether. I have shown you that there is no particular in which plants will not vary from each other; it is quite possible

ble globe, and extirpate or replace the other kinds. That is what is meant by NATURAL SELECTION; that is the kind of argument by which it is perfectly demonstrable that the conditions of existence may play exactly the same part for natural varieties as man does for domesticated varieties. No one doubts at all that particular circumstances may be more favourable for one plant and less so for another, and the moment you admit that, you admit the selective power of nature. Now, although I have been putting a hypothetical case, you must not suppose that I have been reasoning hypothetically. There are plenty of direct experiments which bear o

in the same climate, food, and station;--that every plant has multitudinous animals which prey upon it, and which are its direct opponents; and that these have other animals preying upon them,--that every plant has its indirect helpers in the birds that scatter abroad its seed, and the animals that manure it with their dung;--I say, when these things are con

d as that army was, the struggle must certainly have been a terrible one--every one heeding only himself, and crushing through the ranks and treading down his fellows. The writer of the narrative, who was himself one of those who were fortunate enough to succeed in getting over, and not among the thousands who were left behind or forced into the river, ascribed his escape to the fact that he saw striding onward through the mass a great strong fellow,--one of the French Cuirassiers, who had on a large blue cloak-and he had enough presence of mind to catch and retain a hold of this strong man's cloak. He says, "I caught hold of his cloak, and although he swore at me and cut at and struck me by turns, a

y, which do not hurt the black at all, cut off and destroy the white. Then you see there would have been a selective operation performed; if the white man had risen in that way, he would have been selected out and removed by means of the malaria. Now there really is a very curious case of selection of this sort among pigs, and it is a case of selection of colour too. In the woods of Florida there are a great many pigs, and it is a very curious thing that they are all black, every one of them. Professor Wyman was there some years ago, and on noticing n

ble bees build nests, in which they store their honey and deposit the larv? and eggs. The field mice are amazingly fond of the honey and larv?; therefore, wherever there are plenty of field mice, as in the country, the humble bees are kept down; but in the neighbourhood of towns, the number of cats which prowl about the fields eat up the field mice, and of course the more mice they eat up the less there are to prey upon

d red clover, which are fertilised by the visits of the bees; and they are indirect helpers of t